Board Thread:WARNING ALL USERS False Info\Spoilers/@comment-4148963-20140222014835/@comment-24447558-20140222021706

They were Alphas (a requisite to joining Deucalion's Alpha Pack) and as such their eyes were formerly red. However, they killed innocent people, which is why their eyes are blue now after what Jennifer did "broke" that Alpha part of them (as stated by Deaton in the beginning of 3B.